The story of the in-betweens starts off with a boy who meets three great loves in his journeys and learns how each love might influence him as a person lost in a beautiful yet cruel world. It then follows with the footsteps of a girl called Becky, a fleet-footed angel who traverses onto the stony paths in Essex following the misfortune that befell her mentor Mister Chase. Both protagonists eventually find each other in a time of great turmoil and they would have to find something magical together to fend off the threats of a lifetime.